The communities governing chains and layer-2 rollups.
Arbitrum DAO🟢 ActiveThe community governing the largest Ethereum L2.Cosmos Hub (ATOM)🟢 ActiveThe Cosmos Hub is the first chain built on the Cosmos SDK and the origin of the x/gov module that most Cosmos app-chains still run today.GnosisDAO🟢 ActiveThe DAO behind Gnosis Chain and its ecosystem.Helium (HNT)🟢 ActiveHelium is the canonical DePIN – a decentralized physical infrastructure network – and one of the few DAOs whose token rewards are paid for producing a…Mantle (formerly BitDAO)🟢 ActiveMantle is a DAO built around one of the largest treasuries in crypto, and the clearest example of the mega-treasury archetype in the directory.Optimism Collective🟢 ActiveToken House and Citizens' House – bicameral L2 governance.Orbs Network🟢 ActiveOrbs is a proof-of-stake network whose token holders have governed by signed off-chain vote since 2022, and which spent August 2026 formally constituting…Polkadot (DOT & OpenGov)🟢 ActivePolkadot is a layer-0 network: a central relay chain that provides shared security and interoperability to the parachains connected to it.Polygon (POL)🟢 ActivePolygon began in 2017 as Matic Network, an Ethereum scaling sidechain, and is now a family of Ethereum-aligned chains coordinated under the Polygon 2.0 roadmap.Scroll DAO🟢 ActiveScroll DAO is the governance body of Scroll, an Ethereum layer-2 rollup built on zero-knowledge proofs, in which SCR holders and the delegates they choose vote…Starknet (STRK)🟢 ActiveStarknet is an Ethereum layer-2 built by StarkWare on validity proofs: transactions are executed off-chain and a STARK proof is verified on Ethereum…Syndicate Network Collective (SYND)🟢 ActiveThe Syndicate Network Collective (SNC) governs Syndicate Network, an appchain platform built on programmable onchain sequencers.ZKsync (ZK & ZK Nation)🟢 ActiveZKsync is an Ethereum layer-2 built by Matter Labs on validity (ZK) proofs. ZKsync Era went live on mainnet in March 2023, and the stack has since generalised…